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USGS Open-File Report 94-023

Landscape And Climate Of The Southwestern Russian Plain In The Pliocene

Tanya V. Svetlitskaya
Russian Academy of Sciences, Moscow, Russia
Along the northern coast of the Black sea on the southwestern Russian Plain, interbedded continental and marine sediments cover the entire Pliocene. The fossil floras and faunas of these deposits have been studied in detail, providing the basis for paleoclimatic interpretations and biostratigraphic correlations. This report discusses the most complete sections from this region, concentrating on sites where there are distinct boundaries between stages and horizons and where there are abundant paleontological data from marine mollusks, gastropods, ostracodes and terrestrial mammals and palynological data. The Pliocene of this region is divided into three stages (based on paleontological assemblages) which are (from older to younger) the Pontian, Kimmerian, Kuajlnician. Nine sections were studied from fluvial terraces above the flood-plains of the rivers Byk and, Kuajlnik; from the coastal terrace scarps above Khadzibejsky bay and from the Kertch peninsula of Crimea (Fig. 1).

Figure 1. Pliocene localities on the Southwestern Russian Plain
This figure is available as a GIF, PICT, or TIFF (line-art) image.
Paleomagnetic studies for each of these sections were carried out in Laboratory of the Moscow University following the method of A. Khramov and L. Cholpo (1967). Paleoclimatic reconstructions are based on paleofloristic data from the above mentioned sections studied by S.V. Siabraj and on published materials from adjacent regions (A. Negru (1986), S. Medianik (1985), and N. Shchekina (1979) The methodology used to estimate paleoclimatic conditions on the basis of fossil floras was developed by V. Grichuk (1987), following the concepts of J. Iversen (1941) and modified to be applied to Tertiary floras. In the following text I present paleogeographical reconstructions developed from our paleomagnetic and paleofloristic research, combined with previously published information.

Pontian Stage (5.4-4.7 Ma)

During the Novorossijskoe substage, at the beginning of the Pontian age, the Zanklian transgression united the Black and Caspian Seas with the Mediterranean. By the end of early Pontian time (~5.0 to 4.9 Ma?) the sea retreated southwestward and the greater part of the southwestern Russian Plain was exposed.

Marine faunas indicate that the early Pontian sea was closed and brackish. The mollusks assemblages from these deposits include Unio and Anodonta, while the ostracode fauna includes Caspiella, Bakunella and Leptocythere. From these data Ivchenko (1986) estimated that the mean annual water temperature was near +12° to +15°C (today near the city of Odessa the mean annual water temperature is +9.6°C). In the early Pontian forest-steppe prevailed in the western part of the region under investigation -- pine forests grew in the river valleys, and the northern part was occupied by mixed conifer-broadleaved forests.

By the middle of Novorossijskoe time some aridification had taken place in the east and the role of open vegetation communities increased. Chenopodiaceae were important in these communities and possibly these included halophytes, which may have occupied the recently exposed Black Sea shelf.

In the west such changes of the climate were not registered and forest- steppe vegetation continued to grow. By the end of this period forests dominated in both the east and in the west. These were primarily broad- leaved formations with Ulmus, Zelkova, and Fagus. The climate was similar to the contemporary climate of the southern Crimea coast, but the presence of exotic taxa suggest that it was something warmer than today (January mean temperature ~ +5°C and July ~ +25°C)

The Pontian stage of the history of the Hipparion fauna is poorly represented in the former USSR. Only scattered small occurrences of mammals bones have been found in southern Ukraine. This "Tauric complex" (Korotkevich, 1988) includes the first occurrence of Paracamelus, while typical members of the Hipparion fauna were not present at this time. The increased abundance of ostriches suggests greater availability of open dry areas. However, the presence of certain water-swamp birds and tortoises points to the development of marshlands and warm-water environments, whereas the abundance of Cervus and Proboscidea indicate the presence of wooded vegetation. On the whole, this faunal assemblage reflects moderately warm and relatively dry climate.

Kimmerian Stage (4.7-3.4 Ma)

In the middle Pliocene more land was exposed and sea waters were confined to within the limits of the modern Black Sea and Caspian Sea depressions. The Kimmerian Black Sea basin was closed with brackish water. The marine fauna of this period included Dreissenidae, Cardiidae, endemic gastropods, Viviparus, Melanopsis, Planorbis and the freshwater ostracodes Caspiela and Caspiocypris. The majority of the marine fauna had a boreal character with few Mediterranean taxa. At this time the broad-leaved forests were composed mostly of heat-loving and moderately heat-loving species: Acer, Quercus, Juglans, and less common Castanea, Morus, Carya, Fraxinus, and Tilia. By the end of this period the proportion of steppe elements increased and steppe and forest-steppe landscapes appeared. Paleobotanic data from middle Kimmerian sediments indicate a warmer and more humid climate than previously. During the first half of the Kimmerian, the percentage of subtropical genera in the flora (Pittosporum, Parrotia, Nyssa, Aralia, Magnolia, Staphylea, Taxodium) increased over the levels seen during late Pontian, suggesting that temperatures had increased. The maximum temperature increase occurred during deposition of the sediments of the Kamyshburun horizon (4.2 - 4.0 Ma). Mean January temperature rose by almost 2°C (to 6°C) and mean July temperature rose from 22°C to 23°C.

The Kuchurganian and Moldavian faunal complexes that developed during the Kimmerian time indicate the progressive aridification. The earlier (Kuchurganian) included in its composition the main elements of the Hipparion fauna: Deinotherium, Zygolophodon, Hipparion, and Tapirus, and is largely dominated by forest and forest-steppe species. The Moldavian faunal assemblage reflects the progressive drying of the climate: taxa characteristic of humid environments disappear and inhabitants of open areas such as Paracamelus and Equus (which makes the first appearance here) and more moderate climate appear (Korotkevich, 1986). The climate was apparently humid and moderately warm (January - above +4°C; July ~+22°C).

Kuajlnician Stage (3.4-2.3 Ma)

In the late Pliocene this region was an elevated plain on which the modern river network was formed. the sea basin remained only at the south of the area. It was a closed basin with brackish water , but its salinity was less than in Kimmerian time. The marine fauna was inherited the main features of Kimmerian fauna, but was less diverse. The mollusks Limnocardium, and Dreissena appeared during the Kuajlnician, and gastropods were represented primarily by the fresh-water forms Planorbis, Valvata, and Melanopsis. Ostracodes include the typically fresh-water and brackish-water taxa Cypria kurlaevi and Cyprideis.

Mixed conifer/broad-leaved forests grew along the northern Black Sea coast at the beginning of this time, suggesting more humid conditions than earlier. Acer, Tilia, Fagus, and Quercus grew with more thermophilous trees such as Pterocarya, Zelkova, Fraxinus, Morus, and Juglans. To the east relatively moist steppe vegetation was present. Compared with that of the Kimmerian, the terrestrial vegetation of Kuajlnik time had lower representations of broad-leaved trees and higher levels of steppe taxa. The climate was rather warm and humid, with mean January temperature near +4°C and July mean temperature near +22°C.

A substantial reduction of the forest area occurred at the end of the Kuajlnik time, and among the remaining trees pine and dark-coniferous types (Picea, Abies, Tsuga, etc.) prevailed. The diversity and number of broad-leaved trees and other warmth-loving species decreased. Pollen spectra of Kuajlnik deposits are close in composition to those of the early Pleistocene and only rare pollen grains of Myricaceae, Moraceae, Rutaceae and Taxodium allow us to assign these deposits to the Pliocene. Collectively, these data point to an intensification of aridification and a decrease in temperature.

This is the time of the appearance of the Chaperon faunal assemblage which includes many modern elements. Warmth-loving animals (giraffes, Tapirus, Deinotherium) disappeared by this time and the warm-moderate types (apart from Archidiscodon, Equus, Cervus, Dicerorinus etruscus, etc.) occured more widely (Korotkevich,1988). Fauna data indicate arid climatic conditions and small mammal assemblages indicate steppe vegetation. The climate was apparently dry with moderately warm temperatures (January about +4°C; July +22°C).

Conclusions

In the beginning of the Pliocene forest-steppe vegetation covered most of the southern Russian Plain. By middle Pliocene time temperate broad-leaved forests begin to prevail, and by the late Pliocene increasingly arid and cold conditions led to more open landscapes with steppe and forest-steppe vegetation. The warmest climates of the Pliocene occurred in middle Kimmerian time (~4.2 to 4.0 Ma), but warmer than modern conditions continued until approximately 2.5 Ma.

References


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