One of the most important results of the Trans-Alaska Crustal Transect investigations is the discovery that more than one third of the North American plate in southern Alaska (Chugach Mountains and Copper River basin) consists of tectonically underplated oceanic lithosphere. In southern Alaska, exposed accreted tectonostratigraphic terranes, include, from south to north along the transect, the Prince William, Chugach, Peninsular, and Wrangellia terranes. Chief results from seismic refraction data include the following: (1) A sequence of layers of low and high velocity (5.7–7.8 km/s), more than 10 km thick, dips gently northward, with its top at the surface in the southernmost Chugach terrane and at more than 20-km depth beneath the southern Peninsular terrane. A high-velocity layer at the top of this sequence corresponds to metabasalt in the southernmost Chugach terrane. This sequence is interpreted to be tectonically underplated fragments of the Kula plate and its sedimentary overburden. (2) An intermediate-velocity layer (6.35–6.5 km/s) at 9-km depth beneath both the northern Chugach and southern Peninsular terranes appears to extend without offset across the deep projection of the suture between these two terranes, the Border Ranges fault system. (3) A crustal “root” between depths of 19 and 57 km beneath the northern Peninsular and Wrangellia terranes appears to extend without offset across the deep projection of the suture between these two terranes, the West Fork fault system. This crustal root also appears to abut the tectonically underplated sequence to the south. Unfortunately, in results 2 and 3 the apparent crosscutting and abutting relationships can not be unambiguously resolved. Three scenarios, A, B, and C, have been offered to explain the evolution of this structure. Scenarios A and B differ in interpretation of the depth extent of the northern Chugach and southern Peninsular terranes. In scenario A these terranes extend to only 9-km depth, where they rest on an unknown middle crust, interpreted as a deeper part of the Peninsular terrane. In this scenario, the northern Chugach and southern Peninsular terrenes moved landward as a tectonic wedge, during tectonic underplating in the latest Cretaceous or early Tertiary, and uplifted the upper crust of the Peninsular terrane to the north. In scenario B, the northern Chugach and southern Peninsular terranes extend to 20-km depth. In this scenario, they were compressed but not detached and displaced during accretion. Scenarios A, B, and C differ in interpretation of the deep crustal root that appears to abut the tectonically underplated sequence. Since the Peninsular and Wrangellia terranes were at or below sea level prior to the mid-Cretaceous, the root was probably emplaced in mid-Cretaceous or younger times. In scenario A the root is lower crust of North America that moved southward during tectonic underplating of the Kula plate. In scenario B the root is, in part, tectonically underplated rocks similar to the Kula plate sequence to the south. In scenario C the root is, in part, magmatically underplated rocks.