[1] The Main Ethiopian Rift (MER) is a classic narrow rift that developed in hot, weak lithosphere, not in the initially cold, thick, and strong lithosphere that would be predicted by common models of rift mode formation. Our new 1-D seismic velocity profiles from Rayleigh wave/receiver function joint inversion across the MER and the Ethiopian Plateau indicate that hot lower crust and upper mantle are present throughout the broad region affected by Oligocene flood basalt volcanism, including both the present rift and the adjacent Ethiopian Plateau hundreds of kilometers from the rift valley. The region of hot lithosphere closely corresponds to the region of flood basalt volcanism, and we interpret that the volcanism and thermal perturbation were jointly caused by impingement of the Afar plume head. Across the affected region, Vs is 3.6-3.8 km/s in the lowermost crust and ???4.3 km/s in the uppermost mantle, both ??0.3 km/s lower than in the eastern and western branches of the East African Rift System to the south. We interpret the low Vs in the lower crust and upper mantle as indicative of hot lithosphere with partial melt. Our results lead to a hybrid rift mode, in which the brittle upper crust has developed as a narrow rift along the Neoproterozoic suture between East and West Gondwana, while at depth lithospheric deformation is distributed over the broad region (??400 km wide) thermally perturbed by the broad thermal upwelling associated with the Afar plume head. Development of both the East African Rift System to the south (in cold, strong lithosphere) and the MER to the north (in hot, weak lithosphere) as narrow rifts, despite their vastly different initial thermal states and depth-integrated lithospheric strength, indicates that common models of rift mode formation that focus only on temperature, thickness, and vertical strength profiles do not apply to these classic continental rifts. Instead, inherited structure and associated lithospheric weaknesses are the primary control on the mode of extension. ?? 2009 by the American Geophysical Union.