The interaction of ground water with prairie wetlands in the Cottonwood Lake area has been the focus of research by the U.S. Geological Survey and the U.S. Fish and Wildlife Service since 1977. During this time, climatic conditions at the site ranged from near the driest to near the wettest of the century. Water levels in wetlands and in water-table wells throughout the study area responded to these changing climate conditions in a variety of ways. The topographically highest wetlands recharged ground water whenever they received water from precipitation. The wetland of principal interest, Wetland P1, which is at an intermediate altitude, received ground-water discharge much of the time, but it also had transpiration-induced seepage from it along parts of its perimeter during all but the wettest year. The large fluctuations of the water table in response to recharge and transpiration reflect the ease with which water moves vertically through the fractured till. Lateral movement of ground water is much slower; pore-water moves vertically through the fractured till. Lateral movement of ground water is much slower; pore-water velocities are generally less than 3 m yr⁻¹. The water supply to the wetlands is largely from precipitation during fall, winter, and spring. During these periods, precipitation either falls directly on the wetland, or precipitation that falls on the upland runs over frozen soils or saturated soils into the wetland. The average ratio of stage rise to total overwinter precipitation was 2.59 for the 12-year study period. After plants leaf out, precipitation generally results in much lower rises of the wetland water level. The average ratio of stage rise to over-summer precipitation was less than 1.0.