We elaborate on the well-known spatial association between axc volcanoes and Wadati Benioff zones and explore in detail their genetic relationships as dual physical expressions of slab metamorphism of the oceanic crust and uppermost mantle. At hypocentral depths less than 200 km intra slab Wadati-Benioff earthquakes tend to occur near the top surfaces of slabs. Subduction of very young lithosphere (age < 15-25 Ma) with high heat flow (> 75 mW/m 2) produces mainly shallow earthquakes and spaxse or absent arc volcanism. Subduction of older crust with normal heat flow (50-65 mW/m 2) produces markedly deeper intraslab earthquakes and generally normal volcanic vigor. Seismological observations show that the low-seismic-velocity gabbroic mineralogy of the crust may persist to depths of as much as 150 km in old, cold lithosphere but only to depths of 50-60 km in young, warm slabs. Metamorphic processes in the crust and shallow upper mantle of subducting slabs and the reactivation of faults originally created at shallow depths in the ocean basins probably control the occurrence of intraslab earthquakes to depths of as much as 350 kin. A conceptual model for this metamorphism incorporates the likely effects of water liberated by dehydration. Such dehydration facilitates both brittle faulting by fault reactivation and promotes the kinetics of the transformation of the anhydrous gabbro component of the crust to eclogite. Finite-element modelling shows that densification to eclogite is expected to produce extensional stresses in transformed crust and a smaller compression in the underlying mantle. This model helps explain why most intermediate-depth intraslab earthquakes occur just below the top surfaces of slabs and why many have focal mechanisms indicating down-dip extension. Young, warm slabs have mostly shallow intraslab earthquakes and sparse axc volcanoes because dehydration and eclogite formation largely cease before such slabs axe in contact with asthenosphere. These processes are evidently delayed by kinetic hindrance and the high pressure stability of hydrous phazes at low temperatures in older, colder subducting crust, and thus earthquake activity and asthenospheric-wedge melting tend to be focused at depths of 100-170 kin. Anomalous behavior correlated with the subduction of island-and seamount chains appears to be associated with anomalou shallow intraplate faulting and with perturbations of slab metamorphism by these chains.